Recent Publications

Courrent, A., A. Quennedey, C.A. Nalepa, A. Robert, M. Lenz, and C. Bordereau. 2008. The fine structure of colleterial glands in two cockroaches and three termites, including a detailed study of Cryptocercus punctulatus (Blattaria, Cryptocercidae) and Mastotermes darwiniensis (Isoptera, Mastotermitidae). Arthropod Structure & Development 37: 55-66.

Maekawa, K., T. Matsumoto and C.A. Nalepa. 2008. Social biology of the wood-feeding cockroach genus Salganea (Dictyoptera: Blaberidae: Panesthiinae): did ovoviviparity prevent the evolution of eusociality in the lineage? Insectes Sociaux 55 (2): 107-114, doi 10.1007/s00040-008-0997-2

Farine, J.P., K. Maekawa, K. Shimada, C. Everaerts, P. Luykx, R. Brossut and C.A. Nalepa. 2008. The Cryptocercus punctulatus species complex (Dictyoptera: Cryptocercidae) in the Eastern United States: comparison of cuticular hydrocarbons, chromosome number, and DNA sequences. Molecular Phylogenetics and Evolution 47: 950-959.

Klass, K.-D., C.A. Nalepa and N. Lo. 2008. Wood-feeding cockroaches as models for termite evolution (Insecta: Dictyoptera): Cryptocercus vs. Parasphaeria boleiriana. Molecular Phylogenetics and Evolution 46: 809-817.

Nalepa, C.A. 2007. Harmonia axyridis (Coleoptera: Coccinellidae) in buildings: relationship between body height and crevice size allowing entry. Journal of Economic Entomology 100 (5): 1633-1636.

Nalepa, C.A. 2007. Fly catcher effect in Harmonia axyridis (Pallas)? No attraction to lady beetles images on flight traps. Journal of Entomological Science 42(4): 496-500.

Bell, W. J., L. M. Roth and C. A. Nalepa. 2007. Cockroaches: Ecology, Behavior, and Natural History. The Johns Hopkins University Press, Baltimore, MD, 230 pp.

Lo, N., M.S. Engel, S. Cameron, C.A. Nalepa, G. Tokuda, D. Grimaldi, O. Kitade, K. Krishna, K.-D. Klass, K. Maekawa, T. Miura, and G.J. Thompson. 2007. Save Isoptera. A comment on Inward et al.  Biology Letters 3 (5): 562-563.

Nalepa, C.A. and A. Weir. 2007. Infection of Harmonia axyridis (Coleoptera: Coccinellidae) by Hesperomyces virescens (Ascomycetes: Laboulbeniales): role of mating status and aggregation behavior. Journal of Invertebrate Pathology 94: 196-203.

Lo, N., P. Luykx, R. Santoni, T. Beninati, C. Bandi, M. Casiraghi, W.-H. Lu, E.V. Zakharov, and C.A. Nalepa. 2006. Molecular phylogeny of Cryptocercus wood-roaches based on mitochondrial COII and 16S sequences,and chromosome numbers in Palearctic representatives. Zoological Science 23: 393-398.

Noda, S. T. Inoue, Y. Hongoh, M. Kawai, C.A. Nalepa, C. Vongkaluang, T. Kudo and M. Ohkuma. 2006. Identification and characterization of ectosymbionts of distinct lineages in Bacteroidales attached to flagellated protists in the gut of termites and a wood-feeding cockroach. Environmental Microbiology 8 (1): 11-20.

Nalepa, C.A., G.G. Kennedy and C. Brownie. 2005. Role of visual contrast in the alighting behavior of Harmonia axyridis (Coleoptera: Coccinellidae) at overwintering sites. Environmental Entomology 34(2): 424-431.

Nalepa, C.A. 2005. Cryptocercus punctulatus (Dictyoptera: Cryptocercidae): dispersal events associated with rainfall. Entomologist’s Monthly Magazine 141:95-97.

Nalepa, C.A., G.G. Kennedy and C. Brownie. 2004. Orientation of multicolored Asian lady beetles to buildings. American Entomolologist 50(3):174-175.


Lo, N., C. Bandi, H. Watanabe, C. Nalepa and T. Beninati. 2003. Evidence for cocladogenesis between diverse dictyopteran lineages and their intracellular endosymbionts. Molecular Biology and Evolution 20: 907-13.

Nalepa, C.A.. 2003. Evolution in the genus Cryptocercus (Dictyoptera: Cryptocercidae): no evidence of differential adaptation to hosts or elevation. Biol. J. Linn. Soc. 80: 223-233.

Deitz, L.L.,C.A. Nalepa and K.-D. Klass. 2003. Phylogeny of the Dictyoptera re-examined. Entomologische Abhandlungen 61(1): 69-91.

Perry, J. and C.A. Nalepa. 2003. A new mode of parental care in cockroaches. Insectes Sociaux 50: 245-247.

Nalepa, C.A. and K.A. Kidd. 2002. Parasitism of the multicolored Asian lady beetle (Coleoptera: Coccinellidae) by Strongygaster triangulifer (Diptera: Tachinidae) in North Carolina. J. Entomol. Sci. 37(1):124-127.

Nalepa, C.A., P. Luykx, L.L. Deitz and K.-D. Klass. 2002. Distribution of karyotypes of the Cryptocercus punctulatus species complex (Dictyoptera: Cryptocercidae) in the Southern Appalachians: relation to habitat and history. Ann. Entomol. Soc. Amer. 95(3): 276-287.

Jones, S.C. and C.A. Nalepa. 2002. Survey of the termites (Isoptera: Kalotermitidae, Rhinotermitidae, Termitidae) of St. John, U.S. Virgin Islands. Sociobiology 39(1): 155-163.

Nalepa, C.A., L. Li, W. Lu and J. Lazell. 2001. Rediscovery of the wood-eating cockroach Cryptocercus primarius (Dictyoptera: Cryptocercidae) in China, with notes on ecology and distribution. Acta Zootaxonomica Sinica 26: 184-190.

Nalepa, C.A., L.R. Miller and M. Lenz. 2001. Flight characteristics of Mastotermes darwiniensis. Insectes Sociaux 48: 144-148.

Nalepa, C.A., D. Bignell and C. Bandi. 2001. Detritivory, coprophagy, and the evolution of digestive mutualisms in Dictyoptera. Insectes Sociaux 48: 194-201.

Nalepa, C. A. 2001. Cryptocercus punctulatus (Dictyoptera: Cryptocercidae) from monadnocks in the Piedmont of North Carolina. J. Entomol. Sci. 36: 329-334.

Meyer, J.R., C.A. Nalepa and C. Devorshak. 2001. A new species of Anicetus (Hymenoptera: Encyrtidae) parasitizing terrapin scale [Mesolecanium nigrofasciatum (Pergande)]. Fla. Entomol. 84(4): 686-690.

Nalepa, C.A., K.A. Kidd and D.I. Hopkins. 2000. The multicolored Asian lady beetle (Coleoptera: Coccinellidae): orientation to aggregation sites. J. Entomol. Sci. 35: 150-157.

Nalepa, C.A. 2000. Dampwood termites. In: R.E. Gold and S.C. Jones (eds.). Handbook of Household and Structural Insect Pests. Entomological Society of America, Lanham, MD. Pp. 113-116.

Nalepa, C.A. and M. Lenz. 2000. The ootheca of Mastotermes darwiniensis Froggatt (Isoptera: Mastotermitidae): Homology with cockroach oothecae. Proc. Roy. Soc. London Series B 267: 1809-1813.

Sacchi, L., C.A. Nalepa, M. Lenz, C. Bandi, S. Corona, A Grigolo and E. Bigliardi. 2000. Transovarial transmission of symbiotic bacteria in Mastotermes darwiniensis Froggatt (Isoptera: Mastotermitidae): Ultrastructural aspects and phylogenetic implications. Ann. Entomol. Soc. Amer. 93: 1308-1313.

Nalepa, C.A. and C. Bandi. 2000. Characterizing the ancestors: paedomorphosis and termite evolution. In: Termites: Evolution, Sociality, Symbioses, Ecology. T. Abe, D. E. Bignell and M. Higashi (eds.). Kluwar Academic, Dordrecht, The Netherlands. Pp. 53-75.